The Skull Knight is the boss of the first stage in Dracula's Curse, the town of Warakiya. He can be found in a graveyard at the end of the level, where he first appears only as a pile of old bones. However, when Trevor approaches, he suddenly comes back to life and reassembles himself to confront the hero. His only attack is a quick downward slash, but he can also damage Trevor by touching him, and this is what he'll try to do, by walking steadily toward him, only stopping every once in a while to slash with his sword. He has a considerable amount of HP, but a few flasks of Holy Water should be enough to defeat him in just a matter of seconds.
The Skull Knight reappears in Harmony of Dissonance as the boss of the Skeleton Cave in Castle B. He will change into three forms that are triggered as he sustains damage. His first form has a sword and shield, and attacks by stabbing at Juste. After losing enough HP, he falls to pieces. He regenerates from his shield and drops his sword. Now he attacks mainly by dashing quickly across the room. After losing more HP, he repeats his rebuilding sequence, then drops his shield. In this final form he can shoot a large laser beam from his chest that travels across the entire room.
The Skull Knight is one of the only two bosses in the game which receives non-neutral damage from an element, the Skull Knight being weak to Thunder, and the Golem who, on the contrary, resists to this same element.
It should be noted that in this incarnation, instead of being the remains of a dragon-like creature, he's now the skeleton of a huge warrior who wears a helmet fashioned from what appears to be a bull's skull.